1: Severn Estuary, England. 2: Dee Estuary, England. 3: Menai Strait, Wales. 4: illustration by A. Hancock (in Eliot, 1910). 5: Orkney.
i) No digitiform rhinophores but many have raised ribs/sharp edges to recessed eye-area.
ii) Pale metapodium (tail) absent or negligible.
iii) Often very small, random, light freckles but no distinct, pale, central mark on dorsum.
iv) Pale eye patches.
On saltings with very low salinity, 3‰, to fully marine salinity (though slow to adapt to change in salinity). Sublittoral in inner Baltic.

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