09 Littorina arcana P8560. Shell height 12 mm. Anglesey, Wales. March 2014. Specimen P.

09 Littorina arcana P8560. Shell height 12 mm. Anglesey, Wales. March 2014. Specimen P.

Live male with extended yellowish-white penis; long, slender, steadily tapering to a non-mucronate tip. Not anaesthetised. Inset of same penis at same scale, contracted and distorted by boiling.

Hannaford Ellis (1979) states:
“The range of variation of the penis morphology seen in L. arcana overlaps, to a limited extent, that observed in L. rudis [=L. saxatilis]. This makes it unreliable as a diagnostic feature of either species. Nevertheless it is possible to describe a typical ‘rudis’-shape of penis and a typical ‘arcana’-shape. – – – the penis tip of L. rudis [=L.saxatilis] is frequently mucronate [= with a blunt end and small terminal point]. This never occurs in L. arcana in which the tip tapers to a point. – – – – That it is not always apparent in penes of L.rudis could be due to the killing technique [boiling water in the case of Ellis] affecting the contraction of particular muscles.”
Reid (1996) illustrates penes from live anaesthetised specimens, but they too have their shape altered from that on un-anaesthetised live specimens.

Photographs of penes on live un-anaesthetised animals may prove more reliable, but as they are time consuming to obtain, and the positions the organs are held in varies, it is unlikely that it will become a widely adopted method of identification. See P8554 & P8548

No full account yet of this species.
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