41.1 Cepaea hortensis album. Radula track of unobserved species of gastropod on a pale butyl-rubber roof surface partly covered with algae. Magnified section on right. Oban, Scotland. September 2018. © C. Craik.

41.1 Cepaea hortensis album. Radula track of unobserved species of gastropod on a pale butyl-rubber roof surface partly covered with algae. Magnified section on right. Oban, Scotland. September 2018. © C. Craik.

Observations of radula tracks in the wild (Craik & Anderson, 2018) show that the track of an unobserved gastropod species was linear with few radula strokes per head-swing (1) on surfaces poor in alga not worth expending energy on, and changed immediately (2) on reaching an alga-rich area to head swings five times wider with many more radula strokes per swing (3) and a sinuous route, avoiding previously cleared tracks, to clear a non-linear patch of algae (4).
The radula strokes also removed the weathered grey surface of the butyl to expose paler fresh material.

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